Graptolites     Radiolaria     Stratigraphy

• Origin and Evolution of Animal Coloniality

• Pterobranchia & Graptolithina: Taxonomy and Evolution of the Hemichordata

• Palaeozoic Radiolaria

• Ecology of the marine planktic Ecosystem in the Palaeozoic

• Palaeozoic Chronostratigraphy
   

The evolution of animal coloniality is a complex and astonishing issue with more open-ended questions than solved ones. Colonial organisms like the extinct Graptolithina offered a lifestyle unique to the Palaeozoic, but were doomed through evolutionary restrictions not well understood.  They show a distinctly limited constructional disparity due to their unique skeletal development, preventing them from attaining a leading role, but permitting a limited success in the marine ecosystem. I am interested in micro- and macro-evolutionary patterns of these Hemichordata as a unique example of the evolution of modular colonial organisms in the marine ecosystem.  Phylogenetic interpretations and research on constructional morphology of the colonies may help to understand the needs of these complex organisms for their special planktic life style. Spacial and temporal distribution of faunas might shed light on the factors responsible for origination and extinction events in the evolutionary history of the faunal groups involved and are an important tool to paleogeographic interpretations.

   Isolated Saetograptus sp. cf. S. leintwardinensis,
   Glacial boulder, northern Germany (Maletz)

Radiolarians appear first in the early Palaeozoic, but their occurrence is poorly documented and their phylogeny and evolution nearly unknown. Serious problems in the taxonomy of Paleozoic radiolarian faunas still prevent them from being used successfully for ecological, biogeographical and biostratigraphical interpretations. Newly discovered rich radiolarian faunas of Ordovician age may shed light into the early evolution of their skeleton complexity and to understand the evolutionary lineages leading to modern day radiolarians.

   Radiolarian skeleton development (Maletz)
 

I am interested in the evolution of the marine plankton and its interactions in the Paleozoic; interactions between planktic faunas, the environment, climate and paleogeography to document extinction and origination rates and how they influence evolutionary patterns. The early evolution of planktic organisms is still a mystery, even though planktic organisms are definitely derived from benthic ones, but the exact pathways of their evolution and the driving forces behind it have not been deciphered so far.

   Graptolite bio-sequence stratigraphy at Mt. Hunneberg
   (Egenhoff & Maletz)

I am using a high-resolution chrono- and biostratigraphic correlation approach to get data to trace and specify macro- and microevolutionary patterns and to attain detailed data sets for stratigraphic and phylogenetic research on graptolites. Therefore I am involved in a number of subcommissions of the ICS. It also includes the preparation of the Silurian Monograph of the German Subcommission of Silurian Stratigraphy (eds. T. Heuse, D. Leonhard & J. Maletz). Planktic colonial organisms like the Graptolithina are generally considered as facies independent. Their distribution pattern is based on ecological factors like water mass specifities and temperature gradients. Thus, they are prime suspects for fine-scale stratigraphic correlation on a worldwide scale.

   Radiolarian biostratigraphy (Maletz & Bruton)

My goal is using paleontological data in a geological context, as they essentially represent just one type of information available for valuable geological interpretations, whether they are sedimentological, structural or represent any other aspect of geology.

NSF 0519051

- Graptolite faunal distribution and biogeography in the Middle Ordovician of mid-continent (Great Basin region: Utah, Nevada, Idaho) North America. – Submitted to NSF.

- Graptolite macroevolution: Phylogenetic analysis and testing hypothesis of directional change. – NSF Grant, C.E. Mitchell, J. Maletz (Buffalo, N.Y.), D. Goldman (Dayton, Ohio), in connection with R.A. Cooper (Lower Hutt, New Zealand), P. Sadler (Riverside, Calif.), E.D. Brussa (Salta, Argentina).  

- Chrono- and biostratigraphy in the Paleozoic (Cambrian – Devonian), integrating sedimentological and paleontological methods to achieve enhanced biostratigraphic reliability for geological interpretations (biostratigraphy, paleogeography, structural geology). For this purpose fieldwork is done in a number of regions:

a) Ordovician-Silurian successions in North and South America (New York State, Quebec, Bolivia, Argentina).  Litho- and biostratigraphy, paleogeography, evolutionary studies on graptolites. C.E. Mitchell (UB Buffalo, N.Y), D. Goldman (Dayton, Ohio), R. Suarez-Sorruco (Cochabamba, Bolivia), S.O. Egenhoff (TU Bergakademie Freiberg, Germany).  

b) Paleozoic successions of the Scandinavian platform, a unique intermediate to cold-water carbonate platform without modern examples. Dr. S. O. Egenhoff (TU Bergakademie Freiberg, Germany) is responsible for the carbonate sedimentological investigations. Dr. S.S. Stouge (Copenhagen, Denmark), A.T. Nielsen (Copenhagen, Denmark)

c) Paleozoic successions in Central Europe and their integration in plate tectonic models, mainly focussing on pre-Variscan geology. H. Feldrappe (BGR, Berlin), Prof. Dr. G. Katzung (Berlin). 

Other projects 

Treatise on Invertebrate Paleontology, part V, Graptolithina (eds. C.E. Mitchell, Buffalo, N.Y. & M.J. Melchin, Antigonish, Canada). Responsibility for taxonomy of the Dichograptacea, Glossograptacea.

Revision and updating of the geological aspects (Historical Geology & Time Scale) in the „Brockhaus“-Encyclopaedia for Bibliographisches Institut Gmbh, Gütersloh, Germany, together with Sven O. Egenhoff (TU Bergakademie Freiberg, Germany) and Helga DeWall (Univ. Würzburg, Germany).

Origin and evolution of pterobranchs in the Early to Middle Cambrian– to be submitted to NSF

Earliest pterobranchs were discovered in the Cambrian Wheeler Shale and the Burgess Shale Formation of North America. Similar organisms are present in the Chengjiang biota of China and in the Middle Cambrian succession of the Czech Republic. Their detailed structure and development may shed light on their early evolution and the question of the origin of coloniality in this group of marine organisms. Earliest pterobranch faunas are rarely described and only few specimens have ever been discovered. However, often misunderstanding and even misinterpretation of colonial pterobranchs as fossil algae has hampered the understanding of this group and its evolutionary patterns.

Ordovician Radiolaria in western Newfoundland – Integrated biostratigraphy of the Cow Head Group – to be submitted to NSF

Radiolaria in the Lower to Middle Ordovician Cow Head and Table Head Groups can provide vital biostratigraphic information for correlation and interpretation of early Paleozoic deepwater successions in which other faunal elements are not present. The Cow Head and Table Head Groups are well known from their fossil content and biostratigraphic data are available from diverse faunal groups, including  graptolites, conodonts and chitinozoans. The integration of radiolarian faunas, thus, establishes important ties for a wider application of biostratigraphic time scales for structural geology and even sequence stratigraphic purposes.

DFG (German Science Foundation) Research Grant DFG-Ma 1269/5-1 (July 1999 to June 2001): Graptolite biodiversity in the early Paleozoic: Interactions of ecological factors with evolution and Extinction events.

DFG Habilitation Grant Ma 1269/4-1 (January 1996 to December 1997) : Terrane analysis of the Lower to Midle Ordovician of eastern Avalonia, Baltica and Laurentia using graptolite faunas and their paleogeographic distributions.

DFG Research Grant Ma 1269/2-1 (April 1993 to March 1995): Investigation of proximal end structure of late Arenig to Llanvirnian diplograptid graptolites and their phylogenetic relationships.

DUGW-Projekt (1992 - 2000): Precambrian to Silurian of the southern Baltic Sea: The Adlergrund G14 drill core. incl. Prof. Dr. G. Katzung, Prof. Dr. I. Hinz-Schallreuter, PD. Dr. R. Schallreuter, Dipl.-Geol. H. Beier (Univ. Greifswald)

NATO Project CRG.900816 (1991-1997) M. Vanguestaine (Liege, Belgium), B.-D. Erdtmann (Berlin, Germany), Jörg Maletz (Berlin), Thomas Servais (Liege). Title: Ordovician acritarch/graptolite stratigraphy, and correlation between Belgium and Germany (TU Berlin/Université de Liège)